I haven't noticed any changes in breeding as colonies get larger, but the adult males will kill each other if they don't have enough space to each have a small territory to themselves. For my G. oblongonotas, this space was about a 12" diameter area, but with my A. insignis, they seem happy with about a 5" diameter area. As they're breeding, I'm seeing more and more newly adult males losing legs and antenna as the reigning males hold their territory. The same happened with my G. oblongonota and the king male ended up killing all of the other adult males in the bin. He was particularly feisty, though. I ended up separating him from the colony. That being said, I haven't noticed the same with Elliptorhina, only Gromphadorhina.